GLUScope: A Tool for Analyzing GLU Neurons in Transformer Language Models

We analyze the learned input-output behavior of GLU-based neurons in large language models (LLMs). We propose a simple analysis method: For each neuron, we compute the cosine similarities between its input (reading) and output (writing) weight vectors. In this scheme, a strong negative cosine similarity indicates the neuron weakens the direction it detects in the residual stream, so we call this a weakening neuron. This allows us to gain a number of novel insights. First, we show that nine different LLMs have similar patterns: weakening neurons appear mostly in late layers whereas their counterparts, (conditional) strengthening neurons, are frequent in early-middle layers. Second, we find that weakening neurons display surprising behavior: even though there are few, they activate often and have a large influence on model behavior. Third, weakening neurons have a strong effect on model output when gate values are negative -- which is surprising since negative gate values are not expected to encode functionality.

Biological neural systems employ diverse neurotransmitters -- glutamate, GABA, dopamine, acetylcholine -- to implement distinct signal-processing modalities within shared neural circuits. In contrast, modern transformers apply a single fixed activation function across all feed-forward neurons. We introduce PolyGLU (Polychromatic Gated Linear Unit), a drop-in replacement for SwiGLU that enables each FFN neuron to dynamically route among K=4 activation functions via a differentiable mechanism combining learned static preferences with input-conditioned gating, trained end-to-end with Gumbel-Softmax. We train PolychromaticLM, a 597M-parameter transformer, on ~10B tokens using a single NVIDIA A100 GPU. Our key finding is emergent routing behavior: without any explicit sparsity loss or entropy regularization, the routing mechanism converges to near-deterministic activation selections (mean dynamic entropy = 0.030% of maximum), with a striking depth-dependent specialization pattern -- early layers prefer GELU while deep layers strongly favor Tanh. Three layers maintain elevated routing entropy, suggesting computational flexibility points. The routing architecture adds only 0.23% parameter overhead (~1.4M parameters) and proves fully robust to supervised fine-tuning: routing entropy remains constant at ln(4) throughout 13,067 SFT steps. On standard benchmarks, PolychromaticLM achieves 62-89% of Qwen3-0.6B-Base performance despite training on 3,600x fewer tokens. All code, weights, and training infrastructure are released under Apache 2.0.

Understanding the internal representations of large language models (LLMs) is a central challenge in interpretability research. Existing feature interpretability methods often rely on strong structural assumptions--such as linearity or sparsity--that may not hold in practice. In this work, we introduce InverseScope, an assumption-light and scalable framework for interpreting neural activations via input inversion. Given a target activation, InverseScope characterizes its encoded information by generating natural-language inputs that produce nearby activations, grounding abstract internal states in concrete language. To overcome the prohibitive cost of sampling in high-dimensional activation spaces, we propose a novel control-layer conditioning architecture that substantially improves sample efficiency compared to prior token-prepending approaches. We demonstrate that InverseScope reveals rich geometric structure in LLM representation spaces, including sentence-level linear analogies. The framework scales to state-of-the-art open-source models of up to 14B parameters and generalizes to out-of-distribution inputs, enabling systematic analysis of activation neighborhoods.