Extending large language models to low-resource languages is essential for global accessibility, but training separate models per language is prohibitively expensive. Mixture-of-Experts (MoE) architectures address this by adding sparse language-specific parameters, but determining how many experts each layer needs remains an open question. Current approaches allocate experts based on layer-level similarity, yet language processing exhibits fine-grained specialization at individual neurons. We propose NeuronMoE, a method that analyzes language-specific neurons across all transformer components to guide expert allocation per layer based on empirically measured cross-lingual neuron diversity. Applied to Llama-3.2-3B for low-resource languages (Greek, Turkish, and Hungarian), this approach achieves approximately 40% average parameter reduction while matching the performance of the LayerMoE baseline. We find that low-resource language experts independently develop neuron specialization patterns mirroring the high-resource language, which are concentrated in early and late layers. This reveals potential universal architectural principles in how multilingual models organize linguistic knowledge.
Figures & tables
Figure 1: Expert allocation comparison on Llama-3.2-3B for Greek extension. Cross-lingual neuron diversity (red line) exhibits heterogeneous distribution across layers. NeuronMoE allocates experts following this distribution, achieving 41.7% parameter reduction (49 vs 84 experts) compared to LayerMoE’s uniform allocation strategy.
Figure 2: Overview of NeuronMoE’s core insight and allocation strategy. Top: Cross-lingual neuron diversity exhibits dramatic heterogeneity across layers. For example, Layer 0 contains 342 unique language-specific neurons while Layer 10 contains only 7, demonstrating that different layers have vastly different language-specific processing requirements. Bottom: NeuronMoE allocates experts following this empirical distribution (49 experts total), concentrating capacity where specialization occurs. In contrast, LayerMoE’s similarity-based approach over-allocates in middle layers (84 experts total). This neuron-guided strategy achieves 41.7% parameter reduction while maintaining comparable performance.
ARC
Belebele
HellaSwag
MMLU
Model
#Exp
EN
EL
EN
EL
EN
EL
EN
EL
Llama-3.2-3B (28 layers):
Dense
-
51.11
31.93
74.11
65.22
76.33
43.38
56.45
41.17
Stage 1 (Expert Initialization):
LayerMoE-S1
84
47.27
35.19
73.89
66.33
75.84
52.58
55.40
43.41
NeuronMoE-EN
37 (-56%)
49.15
33.13
73.56
63.67
76.45
49.03
56.89
43.26
Table 1: Performance on Greek extension across Llama-3.2-3B and Qwen-1.5-1.8B. #Exp indicates the number of experts. NeuronMoE achieves 41.7-50.0% parameter reduction with comparable performance, demonstrating cross-architecture generalization.
Layers
LayerMoE
NeuronMoE
0-2 (Early)
5,5,5
6,2,1
3-10 (Early-Mid)
4,3,3,2,2,2,2,2
1,1,1,1,1,1,1,1
11-18 (Mid-Late)
2,2,2,2,2,2,2,3
1,1,1,1,1,1,1,2
19-27 (Late)
3,3,3,3,4,4,4,4,4
1,1,4,1,4,2,2,4,4
Total
84
49
Table 2: Layer-wise expert allocation comparison (Llama-3.2-3B, Greek extension). NeuronMoE achieves 41.7% parameter reduction by concentrating experts in high-diversity layers while minimizing allocation in middle layers.
ARC
Belebele
HellaSwag
MMLU
Language
Model
#Exp
EN
Target
EN
Target
EN
Target
EN
Target
Turkish
Dense
-
51.11
33.56
74.11
59.11
76.33
42.80
56.45
42.35
LayerMoE
84
49.91
36.55
75.33
61.78
76.28
49.60
56.05
43.51
NeuronMoE
50 (-40%)
49.83
34.50
74.44
61.33
76.22
47.83
56.21
43.37
Hungarian
Dense
-
51.11
34.85
74.11
59.67
76.33
44.63
56.45
43.68
LayerMoE
84
49.91
39.73
75.33
64.22
76.28
51.66
56.05
44.86
Table 3: Stage 1 results for Turkish and Hungarian extension on Llama-3.2-3B. NeuronMoE achieves substantial parameter efficiency while successfully extending to additional low-resource languages. All values in %. Results for LayerMoE baseline are from Zhang et al. (2025) .
Layer-Expert
High-AP
Ratio (%)
L0 base_layer
0
0.00
L0 expert_0
0
0.00
L0 expert_1
27
0.14
L0 expert_2
0
0.00
L0 expert_3
61
0.31
L0 expert_4
19
0.10
Table 4: High-AP neuron counts for Greek across experts in selected layers (total neurons per layer/expert: 19,456). Multi-expert layers show distinct specialization patterns, validating allocation decisions.
Figure 3: Heatmap of high-AP neuron ratios for Greek across all experts. Color intensity indicates Greek specialization strength (red: high, blue: low/none). Multi-expert layers (0, 21, 23-27) show distinct specialization patterns across experts, while single-expert middle layers remain largely unspecialized, validating our neuron-guided allocation strategy.
Appendix figures & tables7 assets
Supplementary material from the paper’s appendix.
Appendix
Layers
Unique Neurons (EN+EL)
0-2
342, 79, 11
3-10
18, 15, 11, 8, 7, 8, 8, 7
11-18
6, 9, 14, 9, 18, 81, 20, 19
19-27
28, 38, 232, 51, 208, 114, 111, 238, 223
Total
1,933
Appendix
Table 5: Cross-lingual neuron diversity across layers (Llama-3.2-3B, Greek extension).
L
EN
EL
TR
HU
L
EN
EL
TR
HU
0
322
21
8
8
14
7
2
1
1
1
73
6
5
3
15
10
8
4
6
2
8
3
3
1
16
11
70
10
8
3
11
7
17
6
17
4
16
15
8
4
13
2
0
1
18
10
9
9
15
5
9
2
2
2
19
9
19
22
23
Appendix
Table 6: Complete layer-wise comparison of language-specific neuron counts across English (base Llama-3.2-3B), Greek, Turkish, and Hungarian
Parameter
Value
Optimization:
Optimizer
AdamW
Learning rate
1e-4
Adam β1 , β2
0.9, 0.999
Adam ϵ
1e-8
Weight decay
0.0
Appendix
Table 7: Complete training hyperparameters for all experiments. All models trained with identical settings for fair comparison.
Layers
Greek
Turkish
Hungarian
0-2 (Early)
6,2,1
6,2,1
5,2,1
3-10 (Early-Mid)
1 × 8
1 × 8
1 × 8
11-18 (Mid-Late)
1 × 7,2
1 × 8
1 × 8
19-27 (Late)
1,1,4,1,4, 2,2,4,4
1,1,1,2,2, 4,4,4,4,3
1,1,1,2,1, 3,4,6,3,2
Total
49
50
47
Appendix
Table 8: Layer-wise expert allocation for Turkish and Hungarian on Llama-3.2-3B (28 layers). Both languages show similar allocation patterns to Greek, with concentration in input-proximal and output-proximal layers.
Table 9: Per-layer expert allocation for LayerMoE and NeuronMoE on Qwen-1.5-1.8B (24 layers). LayerMoE uses attention-layer similarity; NeuronMoE uses cross-lingual neuron diversity analysis.
L
Expert
Total
High-AP
Ratio (%)
0
base
19,456
0
0.00
0
exp_0
19,456
0
0.00
0
exp_1
19,456
27
0.14
0
exp_2
19,456
0
0.00
0
exp_3
19,456
61
0.31
0
exp_4
19,456
19
0.10
Appendix
Table 10: Complete expert specialization statistics for Greek. High-AP neurons indicate strong Greek specialization developed during Stage 1 training. Total neurons per layer/expert: 19,456.
Model
Language
ARC
Belebele
HellaSwag
MMLU
Dense
English
51.11
74.11
76.33
56.45
Nepali
24.12
42.44
32.54
33.77
Bengali
26.26
44.78
33.26
33.48
NeuronMoE-S1 (NE)
English
47.44
71.44
75.29
54.71
Nepali
24.89
44.67
33.07
34.63
NeuronMoE-S1 (BN)
English
48.72
75.33
76.01
55.28
Appendix
Table 11: Stage 1 results for Bengali and Nepali extensions on Llama-3.2-3B.
Mixture-of-Experts (MoE) architectures enable efficient model scaling, yet expert routing behavior across underrepresented languages remains poorly understood. We analyze routing dynamics in two architecturally distinct MoE models -- a pure Transformer (Qwen3-30B-A3B) and a hybrid Mamba-Transformer (Nemotron-3-Nano-30B-A3B) -- using Hebrew as a morphologically rich, low-resource testbed. Both pre-trained models exhibit deep-layer routing collapse: usage entropy drops sharply in final layers and tokens concentrate on a narrow expert subset, a pattern largely absent for English. Continual pre-training (CPT) on balanced bilingual data substantially corrects this imbalance, increasing entropy and shifting routing toward shared, language-agnostic experts; supervised fine-tuning (SFT) alone achieves less complete correction. Extending the analysis to Japanese reveals quantitatively consistent collapse signatures, providing cross-linguistic evidence that the phenomenon is a systematic consequence of pre-training underrepresentation rather than any language-intrinsic property. Routing improvements correlate with consistent downstream benchmark gains, positioning routing entropy and expert specialization as principled diagnostics for multilingual capacity in MoE systems.
Mixture-of-Experts (MoE) models exhibit striking performance disparities across languages, yet the internal mechanisms driving these gaps remain poorly understood. In this work, we conduct a systematic analysis of expert routing patterns in MoE models, revealing a phenomenon we term Language Routing Isolation, in which high- and low-resource languages tend to activate largely disjoint expert sets. Through layer-stratified analysis, we further show that routing patterns exhibit a layer-wise convergence-divergence pattern across model depth. Building on these findings, we propose RISE (Routing Isolation-guided Subnetwork Enhancement), a framework that exploits routing isolation to identify and adapt language-specific expert subnetworks. RISE applies a tripartite selection strategy, using specificity scores to identify language-specific experts in shallow and deep layers and overlap scores to select universal experts in middle layers. By training only the selected subnetwork while freezing all other parameters, RISE substantially improves low-resource language performance while preserving capabilities in other languages. Experiments on 10 languages demonstrate that RISE achieves target-language F1 gains of up to 10.85% with minimal cross-lingual degradation.
Kening Zheng, Wei-Chieh Huang, Jiahao Huo +9
University of Illinois Chicago · University of Maryland · HKUST (Guangzhou)
Mixture-of-Experts (MoE) models are widely used to scale language models, yet their expert routing behavior and adaptation in a multilingual setting remain underexplored. In this work, we study multilingual routing dynamics during continual pre-training of an English-centric MoE model on a multilingual corpus, analyzing how expert usage varies across languages. We find that continual multilingual pre-training leads to diffused, language-agnostic routing in early and middle layers, with language specialization primarily emerging in the final layers. We also show that token-level vocabulary overlap between languages plays an important role in how languages are routed. Motivated by these findings, we propose a parameter-efficient adaptation strategy that updates language-specific and shared experts in the final MoE layers. Experiments on MultiBLiMP and Belebele show that our method achieves a strong performance-efficiency trade-off, attaining competitive performance relative to fine-tuning complete final layers, while updating less than 2% of the parameters. Overall, our findings provide insights into where and how language specialization emerges in MoEs during continual pre-training and provide practical insights for low-resource multilingual adaptation. Our code is available at https://github.com/aditi184/moe-routing-adaptation.
Aditi Khandelwal, Marius Mosbach, Verna Dankers +2