Do large language models contain domain-specific parametric shells: concentrated, causally necessary neuron populations whose removal selectively degrades a target domain while sparing others? We apply a uniform causal methodology across two domain granularities, three model families (1.5B to 7B parameters), and eight domains. At the academic subject level, zero neurons exceed 60% domain selectivity across 939,008 combined FFN neurons and causal damage matrices are flat, despite domain identity being linearly decodable above 85% accuracy. At the language and modality level, 0.65--1.14% of neurons exceed 60% selectivity, damage matrices are near-perfectly diagonal (ratios up to 595:1), and shell neuron sets are essentially disjoint (IoU <0.003). Masking code-selective neurons reduces mathematical reasoning accuracy by 16--24 percentage points across all models; masking Spanish or Chinese neurons leaves it at or below random. Shell strength increases monotonically with scale and shells are spatially interleaved in a pattern that precludes group-level selective quantization. Parametric shells form where and only where training data was modular at the token level.
Neuron pruning is widely used to reduce the computational cost and parameter footprint of large language models, yet it remains unclear whether neurons in task-specific models contribute uniformly to task performance. In this work, we provide empirical evidence for the existence and importance of task-specific neurons through a systematic pruning study on language models specialized for mathematical reasoning and code generation. We introduce an activation-based selectivity metric to identify neurons with low contribution to the target task and prune them while preserving target-task accuracy, and compare selective pruning with random pruning. Selective pruning consistently outperforms random pruning, indicating that activation-based selectivity provides a systematic advantage over random pruning. Reverse pruning experiments further show that removing a small subset of highly task-specific neurons (~10%) causes complete performance collapse, suggesting that there exist task specific neurons and critical task information is concentrated in a small portion of the network. In contrast, selective pruning of less critical neurons (~30% - ~35%) reduces accuracy but still preserves significant performance. We also observed consistent reductions in parameters and runtime VRAM usage, along with improved inference throughput as pruning increases. Experiments on both 1.5B and 7B models reveal a robustness threshold around 15-20% pruning, beyond which accuracy loss and generation failures increase sharply. Fine-tuning substantially recovers performance across pruning levels, particularly for aggressively pruned models. These findings provide empirical evidence of neuron specialization in task-specific language models and offer insights into pruning robustness, model redundancy, and post-pruning recoverability.
M. K. Khalidi Siam, Md. Tausif-Ul-Islam, Md. Reshad Romim Khan +5
Multilingual large language models (mLLMs) contain a small fraction of feed-forward neurons that are sensitive to particular languages, commonly termed language-specific neurons. Existing work measures language specificity using the entropy of each neuron's language-wise probabilities of being active, where a neuron is considered active when its activation value is positive. However, this approach may not fully capture the multilingual nature of mLLMs, where language representations are distributional and mutually related. We propose Distribution-aware Language Neuron selection, which leverages pairwise relationships between per-language activation distributions over the full activation range, including negative values. Specifically, we quantify each neuron's language specificity by clustering languages using pairwise overlap coefficients between their activation distributions. Across two mLLMs and two held-out corpora, our identifier more effectively isolates language-specific causal effects, yielding up to 4.9× higher on-target language damage per neuron while preserving off-target language performance.
Large language models (LLMs) display strong comprehensive abilities, yet the internal mechanisms that support these behaviors remain insufficiently understood. In this work, we show that across a wide range of open-weight Transformers, a subset of neurons remains consistently highly activated during inference across tasks of multiple capability dimensions. By probing along the cross-task activation strength, an extremely sparse subset is isolated, whose removal causes a collapse in model behavior, which we term keystone neurons. Our analysis reveals that keystone neurons are a stable and intrinsic neuron subset of the model that is largely established during pretraining. The parameters associated with these neurons are tightly calibrated during the training process, and their precise values are critical for the capabilities of the model. Building on these insights, we propose a supervised fine-tuning approach that updates only keystone neurons, achieving task gains comparable to or even better than full-parameter fine-tuning while better preserving performance in other capability dimensions, despite modifying a much smaller number of parameters.