Multimodal expansion of large language models (LLMs) enables new perceptual capabilities but often compromises the language intelligence acquired during pretraining. In this work, we investigate this phenomenon from the perspective of internal adaptation dynamics and discover that neurons in pretrained LLMs exhibit heterogeneous plasticity during multimodal learning: some neurons are critical for preserving language capabilities, while others are more adaptive to multimodal knowledge. Based on this insight, we propose NeuPAT (Neuron-aware Plasticity Allocation Tuning), a lightweight and architecture-agnostic framework that allocates neuron-wise update constraints during multimodal instruction tuning. NeuPAT uses a small-scale probing stage to estimate neuron adaptation patterns and selectively protects language-sensitive neurons while promoting multimodal adaptation through more plastic neurons. Experiments across diverse LLM families demonstrate that NeuPAT recovers 94.5% of the language capability degradation caused by vanilla tuning on 11 language benchmarks while maintaining comparable multimodal performance, providing an effective approach for capability-preserving multimodal expansion.
Multilingual large language models (mLLMs) contain a small fraction of feed-forward neurons that are sensitive to particular languages, commonly termed language-specific neurons. Existing work measures language specificity using the entropy of each neuron's language-wise probabilities of being active, where a neuron is considered active when its activation value is positive. However, this approach may not fully capture the multilingual nature of mLLMs, where language representations are distributional and mutually related. We propose Distribution-aware Language Neuron selection, which leverages pairwise relationships between per-language activation distributions over the full activation range, including negative values. Specifically, we quantify each neuron's language specificity by clustering languages using pairwise overlap coefficients between their activation distributions. Across two mLLMs and two held-out corpora, our identifier more effectively isolates language-specific causal effects, yielding up to 4.9× higher on-target language damage per neuron while preserving off-target language performance.
Expanding Large Language Models~(LLMs) to new languages is a costly endeavor, demanding extensive Continued Pre-Training~(CPT) and data-intensive alignment. While recent data-free merging techniques attempt to bypass alignment by fusing a multilingual CPT-enhanced model with its instruct counterpart, they are plagued by a critical trade-off: mitigating parameter conflicts to preserve original abilities inevitably dilutes new language acquisition, and vice-versa. To resolve this conflict, we introduce \method, which upcycles a dense model into a Mixture-of-Experts~(MoE) architecture, allocating different experts to different languages. Alignment ability is then transferred by grafting a MoE-expanded parameter delta~(Δpost) to the CPT-enhanced base model, bypassing the complex alignment phase. Experiments demonstrate \method's superiority even against baselines with similar FLOPs or number of parameters; it improves performance on expanded languages while effectively preserving original capabilities. We further show our approach is highly applicable across different models and Post-training deltas.
Large language models (LLMs) display strong comprehensive abilities, yet the internal mechanisms that support these behaviors remain insufficiently understood. In this work, we show that across a wide range of open-weight Transformers, a subset of neurons remains consistently highly activated during inference across tasks of multiple capability dimensions. By probing along the cross-task activation strength, an extremely sparse subset is isolated, whose removal causes a collapse in model behavior, which we term keystone neurons. Our analysis reveals that keystone neurons are a stable and intrinsic neuron subset of the model that is largely established during pretraining. The parameters associated with these neurons are tightly calibrated during the training process, and their precise values are critical for the capabilities of the model. Building on these insights, we propose a supervised fine-tuning approach that updates only keystone neurons, achieving task gains comparable to or even better than full-parameter fine-tuning while better preserving performance in other capability dimensions, despite modifying a much smaller number of parameters.