Transformers and state-space models (SSMs) are the two dominant families of sequence models, and a central open question is how far the analytical knowledge built for transformers transfers to SSMs. We address this through the lens of layer importance which underpins compression, selective fine-tuning, and interpretability across both families. We decompose layer importance into two distinct notions. \emph{Necessity} captures how much the pretrained model depends on a layer's existing contribution, measured by the loss increase from bypassing it. \emph{Plasticity} captures where the model absorbs new information during fine-tuning, measured by the magnitude of task-specific weight updates. Our analysis reveals that the two families behave fundamentally differently: in every evaluated residual transformer up to 14B parameters, Necessity and Plasticity anti-align across depth, whereas in the evaluated Mamba-style SSMs they point to overlapping regions. The sign of this alignment also predicts downstream adaptation behavior. In the evaluated transformers, concentrating updates in the most plastic layers increases catastrophic forgetting, while this tier-dependent effect disappears in the evaluated Mamba-style SSMs.
Selective state space models (SSMs) have recently emerged as a compelling alternative to transformers, combining competitive performance with substantially improved inference efficiency. At each SSM layer, a sequence of hidden states are propagated by a recurrence, mixing information of different tokens. Despite using a different mechanism, this mixing plays a role analogous to attention in transformers. In fact, recent works have shown that the two architectures may be closer than they first appear, as this recurrence admits a formulation akin to linear attention. In transformers, attention is known to drive the tokens to cluster, i.e., to reach consensus, collapsing in the limit to a single direction. Thus, we ask: does the recurrence at the core of SSMs drive the tokens to consensus, as attention does in transformers? To answer this question, we take a dynamical systems perspective on SSMs, modeling the evolution of tokens across layers as an ordinary differential equation. By exploiting input-to-state stability arguments, we establish local exponential stability of the consensus equilibria and characterize their domain of attraction for time-varying weight matrices, a setting not addressed by previous results. We thereby show that the resemblance between SSMs and transformers does run deeper: the recurrence at the core of SSMs aggregates tokens just as attention does. Numerical experiments on a pretrained Mamba-2 model point to the output gate as the component that regulates the extent of this consensus, preventing the tokens from reaching it in full.
João Pedro Silvestre, Álvaro Rodríguez Abella, Paulo Tabuada
The quadratic scaling of Transformer self-attention has driven the adoption of sub-quadratic Selective State Space Models (SSMs) like Mamba, which compress past context into a fixed-size recurrent hidden state. This strict informational bottleneck raises a foundational question for mechanistic interpretability: do SSMs and Transformers learn fundamentally distinct latent representations? In this work, we employ Sparse Autoencoders (SAEs) to conduct a large-scale, feature-level correspondence analysis between Mamba-130m and Pythia-70m over a 10-million token corpus. Contrary to hypotheses predicting widespread architectural divergence, we find no evidence of systematic representational divergence between architectures: across the observed Jaccard distribution, 99.98% of Mamba features cluster toward the upper alignment boundary, providing preliminary feature-level support for the Universality Hypothesis. We further identify and qualitatively characterize this microscopic fraction (0.02%) of diverging features, finding patterns consistent with the hypothesis that the recurrent bottleneck selectively limits the parsing of rigid syntax rather than broad semantic ontology. We demonstrate that while Pythia's unconstrained attention permits the monosemantic decomposition of distinct formatting edge-cases, Mamba is forced to compress unrelated syntactical anomalies into polysemantic "junk drawer" neurons to preserve state capacity. Collectively, these results suggest that architectural routing mechanisms may have negligible impact on core semantic understanding, with representational divergence confined to extreme structural margins.
Rithin Nagaraj, Rupa Laalasa Oruganti, Prerna Subhashchandra Kunder +1
Recent state-space models (SSMs) such as Mamba achieve language modeling performance comparable to transformers despite relying on fundamentally different architectures. This raises an important question: how do these structural differences influence the geometry and functional nature of their internal representations? We study this question through a multi-scale analysis of representations in transformers, SSMs, and hybrid architecture. First, we find that SSMs distribute their representational information evenly across all dimensions, whereas transformer representations are heavily dominated by a single principal direction. By evaluating hybrid architectures, we observe that the representation space becomes increasingly skewed toward a single dominant direction after each attention layer. Next, we explore how the different geometric spread of representations impacts representational capacity through compressibility. Surprisingly, we find that despite their contrasting geometric structures, both architectures exhibit tightly matched effective capacities. We further investigate whether this skewed geometry affects how concepts are encoded. Using rank-constrained probes, we demonstrate that both architectures encode concepts in subspaces of surprisingly similar dimensionality. Furthermore, we demonstrate that the transformers' dominant principal direction does not inherently encode more conceptual information. Finally, we zoom in and examine the alignment between manifolds, either by analyzing representations of specific topics or by looking at the nearest neighborhoods of tokens, and find that they are highly aligned. Ultimately, our analysis suggests that while transformers and SSMs induce different usage of latent space, they display a striking functional convergence at the level of local semantic manifolds.