Olfactory Perception

A robot sent to a named gas leak must preserve gas identity, estimate the source, and navigate to the resulting goal. We present SmellDiffusion, a simulation pipeline that represents species-specific gas zones in an open-vocabulary olfactory scene graph and shares the selected goal between classical and diffusion planners. Its key components are a peak-local geometric gate for selective source correction and diffusion-based, gas-guided trajectory generation. Among 424 unique source-wind configurations in solved flow, 28 have a concentration peak displaced more than 0.5m from the source. A source-independent geometric gate, calibrated only on the training split and evaluated at the observed peak, detects 9 of 10 held-out displacements at 0.64 precision. Gating a precomputed forward-matching correction reduces mean error on the displaced cases from 1.468m to 0.592m (60%), using matching for only 14/204 cases. All-case mean error falls from 0.205m to 0.180m. All planners receive the same scene-graph source estimate as their goal. In a controlled comparison, best-of-ten diffusion achieves mean gas exposure comparable to gas-guided A* (0.0476 versus 0.0455). A single diffusion proposal takes 41.7ms, compared with 72.3ms for gas-guided A*, although best-of-ten sequential sampling increases total runtime. Plain A* also reaches the same goal and remains the fastest and shortest-path method. Six matched Gazebo runs give mean robot-to-source errors of 0.39m for A* and 0.31m for diffusion.

Machine olfaction trains on pooled public descriptor corpora, but whether a shared descriptor word measures the same thing across corpora has not been tested, nor has the ceiling of what any of them can measure. We audit four corpora from Pyrfume. Conditioning on the molecule makes McNemar's test the exact conditional test of the corpus effect. Corpora disagree heterogeneously across descriptors ($I^2 = 80\%$) and non-uniformly with labelling breadth ($z = 17.2$), so no single offset repairs pooling. Median tetrachoric agreement is 0.795 against median $\kappa$ of 0.413: sources largely concur on which molecules deserve a word and differ on how readily they apply it. Of 109 descriptors with an estimable effect, 36 show large differential functioning on the ETS scale. Against a human panel's reliability, Morgan fingerprints with the full RDKit descriptor block reach 32.9% of achievable; adding every label from two merged corpora reaches 33.9%. The gap does not close with model capacity, encoding choice, more molecules, or more words. The missing variance is valence. One pleasantness rating per molecule reaches 54.6% of achievable (57.1% on an independent older instrument). Valence recovered from descriptors ($\rho = 0.457$) yields only 16.6%, so it must be measured. Five raters exceed structure plus the full descriptor record; fifteen to twenty saturate. We release a descriptor crosswalk and twenty machine-checked theorems.

Olfaction played an indispensable role throughout human evolution and civilization. Even in the contemporary era of advanced technology, olfaction remains a critical channel for person to conduct danger discrimination, emotional experience, and memory formation. However, most substances in nature exist as multi-molecule mixtures. The complexity of mixture compositions, as well as concentration dependent saturation effects and receptor specific activation thresholds, pose substantial challenges in identifying olfactory characteristics. In this study, we proposed a novel bio inspired deep learning framework for accurate odor perception recognition of mixtures. We robustly constructed neural response curves for molecule-receptor interactions, and developed a fusion strategy that integrates attention-weighted multi-receptor curves with concentration-dependent multi-molecule curves, replicating the competitive activation and synergistic integration of mixture components. Furthermore, by comparing the consistency of response curve patterns, the model can transfer knowledge from the semantically rich space of molecular associations to guide recognition of mixture perception characteristics. Therefore, we established a complete computational pathway from chemical blending, neural encoding, to perceptual formation. Finally, we conducted comprehensive evaluation, and results demonstrated exceptional superiority, achieving an accuracy of 92.2%. Consequently, our work provides a generalizable solution to the long standing mixture perception challenge. More importantly, it can be integrated into embodied cognitive systems to enhance the agents perceptual and interactive capabilities in complex scenarios.

Odor quality is commonly represented using high dimensional descriptor profiles, yet their low dimensional organization remains unclear. We investigated whether a two-dimensional hyperbolic embedding can provide an interpretable representation of this structure. We applied hyperbolic metric multidimensional scaling to two complementary datasets: 480 Sagar rating profiles from three participants rating 160 odorants on 15 continuous descriptors, and 4983 GoodScents--Leffingwell molecules annotated with 138 binary descriptors. The embeddings substantially preserved pairwise descriptor distances, supporting subsequent analyses of radial and angular organization. In Sagar, rating profile entropy was strongly and negatively associated with hyperbolic radius, with diffuse profiles closer to the center and concentrated profiles closer to the boundary. This radial organization emerged primarily at the level of the full descriptor profile, rather than any individual descriptor, and remained robust across alternative descriptor representations, participant specific analyses, and averaged ratings. Sweet, musky, fruity, pleasantness showed the strongest directional trends. In GoodScents--Leffingwell, active label entropy, reflecting descriptor multiplicity, increased with radius, whereas orthogonalized descriptor entropy, reflecting spread across orthogonal modes, decreased with radius. Related binary descriptors occupied coherent localized high-density regions. These findings reveal complementary radial and angular organization in the hyperbolic representation of olfactory descriptor data. They support hyperbolic mapping as an interpretable descriptive framework in which radius summarizes global profile properties, while the angular component captures continuous descriptor gradients and categorical organization.

Pretrained molecular encoders are commonly evaluated through downstream prediction, but predictive accuracy alone does not establish that a learned representation captures reproducible scientific structure, adds information beyond strong conventional baselines, or transfers out of distribution. We present a reliability-aware audit of generic molecular representations for human olfaction across four distinct claims: global perceptual geometry, incremental predictive value beyond chemistry, cross-dataset replication, and mixture transfer to unseen components. Using the Keller-Vosshall and Bierling single-molecule rating datasets and the Ma binary-mixture dataset, we compare MoLFormer and ChemBERTa against RDKit descriptors and Morgan fingerprints under identity-controlled and matched evaluations. Human three-attribute rating geometry, based on intensity, pleasantness, and familiarity, is reproducible across participant splits (median RSA 0.743 and 0.855), whereas model-human alignment is substantially weaker (RSA 0.019-0.158). Learned embeddings do not consistently outperform conventional representations in global alignment, and MoLFormer provides no clear incremental predictive value beyond a combined RDKit-Morgan baseline in either single-molecule dataset. Human geometry shows positive but incomplete agreement across 63 shared molecules (RSA 0.331; 95% bootstrap interval [0.204, 0.507]). Under one strict unseen-component mixture split, incremental effects are outcome- and representation-dependent, with all intervals crossing zero. These results establish empirical boundaries for the evaluated generic molecular encoders and motivate a broader evaluation principle: representation quality in scientific domains should be assessed separately for target reliability, structural alignment, incremental information, replication, and out-of-distribution transfer.

Olfaction is important for emotion regulation because it acts as a non-intrusive and cognitively lightweight pathway that directly engages the brain s affective circuitry and achieves unobtrusive emotional modulation. This trait is essential for advancing practical affective computing in daily and attention-critical scenarios. However, current olfactory emotion research has two key limitations. First, it overemphasises the valence dimension while neglecting arousal. Second, it lacks multimodal datasets that synchronously capture central and peripheral physiological responses to olfactory stimuli. To address these issues, we construct a large-scale multimodal olfactory emotion dataset based on 111 subjects, in which odors are labeled in the 2D arousal-valence space and electroencephalogram (EEG), electrocardiogram (ECG), and photoplethysmography (PPG) signals synchronously recorded. Nevertheless, multimodal signals present challenges such as non-stationarity, differences in latency, and cross-modal heterogeneity. Thus, we propose a spatiotemporal-frequency hybrid fusion network (STF-HFNet), which integrates three core modules. Frequency aggregation processing learns adaptive frequency aggregation in order to model non-stationary dynamics. Reciprocal guided attention enables reciprocal bidirectional calibration for cross-modal temporal alignment without synchronisation priors. Hybrid collaborative fusion combines spatial and channel attention mechanisms to enhance cross-modal complementarity while suppressing redundant information. Extensive experiments show that STF-HFNet achieves state-of-the-art (SOTA) recognition accuracies of 88.34% on the AMIGOS dataset and 92.40% on our self-constructed dataset, and outperform the SOTA methods by 8.27% and 5.07%, respectively.

Machine olfaction for gas unmixing is an underconstrained inverse problem in which gas compositions must be inferred from low-dimensional, delayed, and entangled sensor responses produced by interacting chemical transport, surface adsorption, and sensor transduction. One of the key obstacles is physics closure misspecification, where a neural network is designed to fit sensor traces rather than infer a physically closed olfactory process. In this work, we formulate gas unmixing as a multi-physics-constrained inverse problem governed by Maxwell--Stefan multicomponent transport PDEs, competitive adsorption ODEs, and nonlinear sensor transduction ODEs. Directly solving such a high-dimensional coupled system is computationally expensive and often numerically unstable. To this end, we propose UnMixNet, a physics-closed graph neural solver that embeds this multi-physics forward process into end-to-end gas unmixing. UnMixNet discretizes Maxwell--Stefan cross-diffusion on spatial graphs and formulates the multicomponent flux on each edge. This design enables local, differentiable, and flux-conservative inference for multicomponent cross-diffusion. Evaluations on SmellNet show improved single-odor recognition, seen-mixture unmixing, and unseen-mixture generalization. In addition, an external validation on UCI Dynamic Gas Mixtures shows that the inferred concentration process agrees with ground truth concentration set points under dynamic transitions. Process-consistency diagnostics further show that the proposed model learns transferable dynamic physical fingerprints that better satisfies transport, conservation, adsorption, and readout closure.

The Contrastive Olfaction-Language-Image Pre-training 2 (COLIP-2) model is a multimodal embeddings space that places olfaction as a first-class citizen among vision and language. Molecular structure, gas-sensor readings, odor-descriptor language, and images are all trained into a single shared representation space, so that a robot can localize a detected aroma to objects in a scene probabilistically. No ImageNet-scale datasets of paired image-scent examples exists which warrants the need for their collection. Our intent with the release of COLIP-2 is to demonstrate the limit of what can be built for robotics with open-sourced olfactory data in order to ground the argument for why new methodologies and datasets are necessary in order to enable advanced olfactory-oriented perception capabilities. We enumerate results from internal testing of the COLIP-2 architecture and make necessary optimizations to run the model at the edge for real-time robotics applications. While developed with robotics in mind, the design of COLIP-2 has been influenced by experts across many disciplines of science in academia and industry, and we hope that the model can be useful in any multimodal domain requiring olfactory intelligence.

Chemical pollutants released into the environment are transported by turbulent flows, generating complex, intermittent plume structures that threaten ecosystems and human health. Rapid localisation of emission sources is critical, and field robots equipped with chemical sensors provide a viable means to perform this task. However, inferring source location from sensor readings remains difficult due to sparse detections and the absence of reliable concentration gradients. Existing approaches fall into two paradigms. Bio-inspired strategies rely on reactive behaviours triggered by detections, such as surge-casting, offering efficiency but requiring scenario-specific tuning. Cognitive strategies integrate observations into a probabilistic belief over source location. While more robust, they suffer from excessive exploration and strong dependence on belief accuracy. The Fast-Cognitive algorithm reduced this computational burden but preserved the fundamental limitations. Previous Markov chain analysis revealed that source-directed motions occur roughly twice as often following odour detections, indicating that reactive behaviours naturally emerge within cognitive frameworks. This work proposes a hybrid strategy that explicitly incorporates bio-inspired reactivity into belief-dependent motion planning. It introduces a detection-triggered switching mechanism formalising transitions between crossflow exploration and source-directed motion, prioritising source proximity over information gain. Behavioural parameters are derived directly from belief metrics, enabling adaptive reactivity without manual tuning. The approach is validated through simulations under three turbulence conditions and field experiments with an autonomous surface vehicle in the Mondego River, Portugal. Results show up to 50% reduction in travelled distance, 86% success rate, and 3.2m average localisation error.

Images tell us what a scene looks like, but rarely what it would feel like to be there. While recent datasets pair visual scenes with electronic-nose measurements, aligning smell signals with images remains challenging because many olfactory cues arise from contextual environmental factors that are not directly visible in pixels. We introduce SCENT, a multimodal framework that uses language guidance as a semantic bridge between vision and olfaction. Our approach leverages Vision-Language Models (VLMs) to generate scene descriptors capturing objects, environmental context, and plausible ambient smell cues suggested by the visual scene. These descriptors provide semantic guidance for learning olfactory representations. We train a smell encoder that maps electronic-nose signals into a shared embedding space aligned with both visual and textual representations, and introduce a languageguided latent decomposition that separates object-specific odors from contextual environmental contributions. Experiments on the New York Smells dataset demonstrate that SCENT significantly improves crossmodal retrieval compared to vision-only baselines, achieving state-of-theart performance on smell-to-image and smell-to-text retrieval tasks. In addition, our framework produces interpretable olfactory representations that enable the disentanglement of complex smell mixtures. Our results reveal the importance of contextual semantic information for grounding olfactory perception in multimodal learning and pave the way for future research in this area.

Crossmodal correspondences between sound and taste are well established in psychology and neuroscience, but largely absent from content-based multimedia retrieval. We formalise taste-from-audio prediction as a content-based music information retrieval benchmark over a perceptually validated multi-source corpus, comparing ten frozen audio encoders from the four HEAR families under a shared multi-task regression head, with gated late-fusion as a configurable variant. In order to assess the effectiveness of the models, we compute absolute error and rank correlation. The strongest systems predict the five tastes within a macro RMSE of 0.134; on held-out real music their error is less than half a single rater's deviation from the consensus (RMSE 0.13 vs. 0.28), so the model tracks the group consensus more closely than an average human rater, and well below the previous state of the art baseline (0.219). On absolute error the encoders are statistically flat, with a single VGGish matching the best fusion, but gated late-fusion's advantage is confined to rank correlation (macro Pearson r 0.724 vs. 0.666). Operationalised as a content-based retrieval index, the predicted taste space ranks a 309-item pool far more faithfully than a CLAP-text baseline, which sits at chance; ridge probes and an audio-bandstop knockout read the strongest representations against documented sound-taste correspondences.

Inspired by advances in natural language processing and computer vision, "time-series foundation models" (TSFMs) have recently been introduced with the promise of strong generalization across diverse time-series tasks, including forecasting, classification, and anomaly detection, as well as across domains such as healthcare, climate science, and manufacturing. However, their utility for gas-sensing data remains largely unexplored. To address this gap, this paper systematically evaluates recent TSFMs on electronic nose (E-Nose) data. In particular, we investigate whether embeddings produced by representative TSFMs, including Chronos-2 and MOMENT, provide effective representations for gas identification and concentration prediction. Specifically, we show that fine-tuning is necessary to achieve satisfactory performance on E-Nose data, and fusing TSFM embeddings with representations learned by specialized predictive models can further improve the performance, suggesting both the potential and limitations of current TSFMs for gas-sensing applications.

While modern multimodal models integrate vision with language, audio, or touch, olfaction remains largely unexplored due to the lack of paired visuo-olfactory data. We introduce SmellNet-V, a scalable visuo-olfactory dataset built on the insight that odor identity is largely invariant to visual transformations within a semantic category. This allows us to synthetically pair smell-only samples with semantically aligned in-the-wild web images, converting a unimodal olfactory dataset into a cross-modal benchmark without costly co-collection. Building on this dataset, we propose See & Sniff, a self-supervised framework that learns joint visuo-olfactory representations via dense local alignment and naturally produces smell saliency maps for spatial grounding of odor sources. We further introduce pixel-level smell localization task and a benchmark for evaluation. Our method surpasses smell-only baselines by 7% in smell classification from smell alone and generalizes to cross-modal retrieval and smell localization, establishing visuo-olfactory learning as a new direction in multimodal perception.

Named Entity Recognition (NER) in low-resource languages suffers from limited supervision and a lack of high-quality pretrained embeddings. Biological olfaction, which relies on sparse combinatorial coding through receptor and glomerular organization, offers a compelling paradigm for learning robust representations under uncertainty. In this paper, we introduce a receptor-glomerular bottleneck - a novel, biologically-inspired olfactory architecture - between standard token embeddings and a BiLSTM-CRF sequence model. We evaluate our architecture across six multilingual datasets trained entirely from scratch (without pre-trained embeddings) under varied data-scale conditions, including a strict 1k-sentence low-resource control. Our results demonstrate that introducing a representation bottleneck yields F1 score improvements under severe data scarcity, primarily by acting as a powerful regularizer. Under the 1k capped training condition, at least one olfactory-inspired configuration achieves the highest mean F1 score across all six datasets. While these improvements represent near-ties with generic bottleneck controls for most languages, the olfactory architecture provides a significant advantage in languages like Bangla (+6.23% F1 over standard baseline and +8.47% F1 over the best control baseline) where generic bottlenecks degrade performance. We also observe improvements in the ultra-low-resource Telugu setting (+4.43% F1) at full-scale, and find that sparse specialization naturally emerges within the receptor layer. Our findings suggest that structured sparse coding inspired by olfactory networks serves as an effective inductive bias and regularizer when representations must be learned from limited or noisy supervision.

Predicting human olfactory perception from molecular structure has seen remarkable progress, yet these approaches require explicit chemical structure at inference, which is not available in practical sensing settings. We address this gap by exploring direct electron ionization mass spectrometry (EI-MS), a sensing technique that acquires chemically informative fragmentation fingerprints in seconds, as an alternative input modality for olfactory prediction. We contribute Spectrum-to-Chemical Embedding alignmeNT (SCENT), a multi-modal contrastive learning framework that aligns EI-MS representations with pretrained chemical structure embeddings, while requiring only mass spectra at inference. On the multi-label odor descriptor prediction task, SCENT significantly outperforms MS-only baselines and achieves performance comparable to structure-based models, despite requiring no explicit molecular structure at test time. The learned representations also better approximate continuous human perceptual ratings and generalize to real-world lab-measured spectra, suggesting that cross-modal alignment is an effective strategy for grounding analytical spectra in chemical semantics.

Training data for olfaction is scattered through disparate, non-standardized datasets that limit the ability to build representative world models. Olfactory navigation is a highly dynamic and non-stationary task that benefits from real-time continual learning. We introduce an adaptive framework called Grow-Prune-Freeze (GPF) networks that enable an agent to continually learn through growing, pruning, and freezing early layers of its policy in response to world complexity. Grounding GPFs in non-linear random matrix theory, we show that the work of Pennington & Worth (2017) can be extended from single hidden layers to n-layer continual-learning models, and that eigenvalue composition of network weights is preserved as successive layers are added. We show that GPFs based on Expected SARSA achieve a 94% success rate on turbulent plume navigation - a partially observable, non-stationary task representative of the "big world" challenges that motivate adaptive learning in robotics - and provide supporting methodology for applying GPFs in other world models. Further experiments amount evidence that GPFs may generalize well to other machine learning tasks such as reinforcement learning in Atari, image classification, and autoregressive language models. We open source all code and data to encourage improvements on and more research in olfactory robotics.

In dynamic flow fields, various animals exhibit remarkable odor search capabilities despite relying on stochastic detections. Interestingly, there exists an optimal time window for integrating these detections that maximizes search efficiency. To understand the underlying mechanism, we investigate the navigation performance of Reinforcement Learning (RL) agents in unsteady flows under varying memory lengths and flow conditions. Without any predefined models, the agents develop a flow-assisted casting strategy and adaptively adjust both the geometry of their search trajectories and the concentration threshold for initiating casting to maximize the success rate. The agent's average speed toward the odor source exhibits a non-monotonic dependence on memory length, which can be explained by the "sector-search" model.

Finding an odor source in a turbulent flow requires effectively leveraging the history of olfactory observations into a robust navigation strategy. In this work, we use tabular Q-learning to train an olfactory search agent with a minimal memory of past observations: only a running clock since the last whiff. This agent learns an interpretable strategy to recover the plume which combines well-known behaviors observed in insects: surging, casting, and a return downwind. While achieving good performance on data from direct numerical simulations of turbulence, the agent is limited by an inability to adapt its strategy to the local intermittency level; we show that providing more flexibility improves robustness.

Architectural code smells erode software maintainability and are costly to repair manually, yet unlike localized bugs, they require cross-module reasoning about design intent that challenges both developers and automated tools. While large language model agents excel at bug fixing and code-level refactoring, their ability to repair architectural code smells remains unexplored. We present the first empirical evaluation of LLM agents on architectural code smell repair. We contribute SmellBench, a task orchestration framework that incorporates smell-type-specific optimized prompts and supports iterative multi-step execution, together with a scoring methodology that separately evaluates repair effectiveness, false positive identification, and net codebase impact. We evaluate 11 agent configurations from four model families (GPT, Claude, Gemini, Mistral) on 65 hard-severity architectural smells detected by PyExamine in the Python project scikit-learn, validated against expert judgments. Expert validation reveals that 63.1% of detected smells are false positives, while the best agent achieves a 47.7% resolution rate. Agents identify false positives with up to $κ= 0.94$ expert agreement, but repair aggressiveness and net codebase quality are inversely related: the most aggressive agent introduces 140 new smells. These findings expose a gap between current LLM capabilities in localized code transformations and the architectural understanding needed for cross-module refactoring. SmellBench provides reusable infrastructure for tracking progress on this underexplored dimension of automated software engineering. We release our code and data at https://doi.org/10.5281/zenodo.19247588.

The promise of Large Language Models in automated software engineering is often measured by functional correctness, overlooking the critical issue of long term maintainability. This paper presents a systematic audit of technical debt in AI-generated software, revealing that AI does not eliminate flaws but rather introduces a distinct machine signature of defects. Our multi-scale analysis, spanning single-file algorithmic tasks and complex, agent generated systems, identifies a fundamental Reasoning-Complexity Trade-off: as models become more capable, they generate increasingly bloated and coupled code. This architectural decay is so pronounced that we establish a Volume-Quality Inverse Law, where code volume is a near perfect predictor of structural degradation. Crucially, we demonstrate that neither functional correctness nor detailed prompting mitigates this decay. These findings challenge the current paradigm of prompt-driven generation, reframing the central problem of AI-based software engineering from one of code generation to one of architectural complexity management. We conclude that future progress depends on equipping agents with explicit architectural foresight to ensure the software they build is not just functional, but also maintainable.

Olfaction offers robots access to chemical information that is largely inaccessible to vision, touch, and audition, yet using airborne chemical signals for spatial perception remains challenging because local volatile organic compound (VOC) measurements are shaped by complex chemical transport and sensor dynamics. We introduce Scensory, a robotic olfaction framework that learns to jointly infer biological source identity and relative location from short temporal VOC measurements. Using a robot-automated data collection platform, we pair VOC dynamics from cross-sensitive gas sensor arrays with spatial supervision and train models to predict fungal identity, source direction, and distance. We show that a single sensor array can extract all three quantities from only 3 s of local measurements under ambient environmental conditions, achieving species classification accuracy of up to 80.13%, directional accuracy of up to 68.65%, and mean absolute distance errors of 0.110-0.131 m. Incorporating measurements from multiple spatial locations further reduces ambiguity, improving peak species and directional accuracies by 9.72 and 18.66 percentage points, respectively. We then embody this learned olfactory perception on a mobile robot, where successive local predictions acquired during motion are transformed into a world-frame evidence map, allowing observations from different positions and headings to reinforce persistent source hypotheses and guide closed-loop localization. Across eight selected indoor runs, the robot achieves a planar endpoint error of 0.606 +/- 0.294 m. Our results establish airborne chemical dynamics as a viable perceptual signal for robots to recognize biological sources, reason about their spatial origin, and autonomously navigate toward them under ambient environments.

Traditional honeypots share an invariant behavioral profile: a skilled adversary can confirm the presence of a deception environment within a few diagnostic commands, limiting their intelligence value. Commercial deception products (USD 100,000-150,000/year) similarly lack real-time model-driven feedback. Chameleon, an openly distributed adaptive honeypot, addresses both shortcomings. It integrates: a BiLSTM classifier achieving 99.61% accuracy across seven threat categories at ~2 ms CPU latency; a locally deployed Qwen3.5-0.8B model delivering 90% generation accuracy at 4.5 ms latency; and two meta-heuristic engines. Threat-Calibrated PSO (TC-PSO) reshapes swarm inertia and objective amplification in proportion to the classifier's anomaly output, adjusting connection-holding delays in real time. Semantic Deception RRT (S-RRT) evolves deception schemas via exponentially scaled pheromone updates from a language-model severity assessment, with a depth-decay multiplier enforcing a finite memory footprint. A controlled 30-seed benchmark (42-71, identical trajectories and budgets) shows threat-calibrated inertia alone does not improve search over standard PSO on static or dynamic landscapes (p = 0.18); population-diversity mechanisms (GA/ACO) significantly outperform PSO-family optimizers on threat-regime shifts (p < 0.0001, d <= -37). S-RRT's depth-decay delivers a significant memory reduction versus standard RRT (53.1 vs. 119.2 units, p < 0.0001, d = -10.0); its severity-weighted pheromone does not improve raw fitness. Operating cost is ~USD 17/month, a ~490-fold reduction versus commercial alternatives.