Spiking Neural Networks

Deep learning has greatly advanced automatic speech recognition (ASR), enabling widespread deployment on edge devices such as smartphones and smart home systems. However, the computational and energy demands of deep neural networks pose significant challenges for such resource-constrained deployments, introducing latency and limiting real-time interaction. Neuromorphic computing offers a promising solution by introducing activation sparsity through spiking neural networks (SNNs) and event-driven neural networks, converting dense operations into sparse computations. However, a study that evaluates the hardware benefits of different neuromorphic strategies remains lacking for ASR. This paper explores spiking and event-driven neuromorphic neural networks to improve activation sparsity in the state-of-the-art SpeechMamba model for ASR. We introduce an event-driven SpeechMamba with FATReLU activation, achieving over 60% activation sparsity with less than 1% accuracy degradation on LibriSpeech. We also propose a spiking SpeechMamba that attains over 70% sparsity while using 30% fewer parameters than comparable SNNs. Finally, we develop a cycle-accurate event-driven simulator enabling flexible algorithm-hardware co-exploration, which helps us identify computational bottlenecks and yields over 10% additional efficiency improvements.

Spiking neural networks (SNNs) provide event-driven and low-power computation inspired by biological neural systems, but current implementations rely on von Neumann graphics processing units (GPUs) and central processing units (CPUs) platforms, where memory and computation bottlenecks limit energy efficiency. To address this challenge, this paper proposes an analog memristor-based spiking neural network (SNN) accelerator that integrates in-memory synaptic computation with analog integrate-and-fire (IF) neurons, eliminating multi-transistor CMOS synapse circuits and enabling asynchronous event-driven operation at the 45nm technology node. Additionally, a digital SNN accelerator is designed and optimized at the 5 nm technology node for comparison. The proposed architecture is evaluated using a predator-prey tracking task that emulates pursuit behavior. In this task, the analog SNN accelerator's inference closely matches the ideal software inference with a mean squared error (MSE) of 0.004. HSPICE simulation results show that the proposed analog SNN accelerator achieves 12.7 times lower energy consumption and 1.26 times lower delay compared to the digital baseline, demonstrating the potential of memristor-based neuromorphic circuits for energy-efficient real-time edge intelligence.

Training deep spiking neural networks (SNNs) remains challenging due to sharp loss landscapes and temporal inconsistency caused by surrogate gradients. To address these challenges, we propose a unified framework: adaptive and asymmetric surrogate gradients A2SG. The adaptive gradients adjust an effective window for spatio-temporal adaptation, reducing spatial gradient variation and maintaining directional consistency of gradients over time. The asymmetric gradients reflect neuronal dynamics by assigning larger gradients to neurons with higher membrane potentials, and we prove that they yield lower variation than symmetric surrogates. Our analysis further establishes a direct connection between local gradient variation and the curvature of the loss landscape, providing a principled explanation for how A2SG promotes convergence to flatter minima and improves generalization. We conduct extensive experiments on diverse models, including CNN-based and Transformer-based SNNs, across various tasks such as image classification using both static and neuromorphic datasets, as well as segmentation. The results demonstrate that A2SG consistently improves accuracy and energy efficiency, establishing it as a general and reliable solution for training deep SNNs. Our code is available at https://github.com/KIST-NCL/A2SG.git.

This paper proposes SpikeWFM, a novel hybrid architecture that integrates spiking neural networks (SNNs) with conventional artificial neural network (ANN)-based transformers for wireless foundation models (WFMs). Inspired by the noise-robust and energy-efficient information processing in the human brain, SpikeWFM aims to enhance the resilience of WFMs against noise and interference while maintaining strong generalization capabilities across diverse wireless scenarios. Drawing from the success of large language models, WFMs leverage self-supervised pre-training on large-scale datasets spanning various wireless environments to learn a unified embedding that supports a wide range of downstream tasks, including channel prediction, channel estimation, beam predition, positioning and etc. Such models typically outperform task-specific designs and exhibit superior adaptability to unseen conditions. However, existing WFMs remain vulnerable to realistic noise and interference in practical wireless systems. To address this limitation, we incorporate spiking neurons into the transformer-based WFM architecture. We provide a brief theoretical analysis demonstrating how the SNN-ANN hybrid effectively mitigates noise and interference through temporal sparsity and event-driven processing. Experimental results show that SpikeWFM consistently outperforms conventional ANN-based WFMs in both pre-training convergence and channel prediction accuracy. Additional results on communication and sensing tasks will be presented in the full journal version of this work.

Recognizing and continuously learning novel human actions without forgetting prior classes is a requirement for emerging AR/VR and robotics applications. For these applications, both on-device processing and learning are essential for privacy and low-latency adaptation. Event cameras address the efficiency of visual sensing with sparse, asynchronous output that is naturally compatible with neuromorphic processing. Yet no prior system has deployed a continual on-device learning pipeline for event-based action recognition using neuromorphic hardware. We present CLANE, Continual Learning of Actions on Neuromorphic Hardware from Event Cameras, deployed end-to-end on Intel Loihi 2. CLANE combines a spiking 2D CNN for spatiotemporal feature extraction with CLP-SNN as its on-chip learning head, extended to action clips via a Temporal Aggregation Layer and a fixed-point Normalization Layer, both novel Loihi 2 modules. On THU E-ACT-50, a 50-class dataset captured under real-world conditions, CLANE achieves 70.4% accuracy in a continual learning task while delivering more than 100x energy reduction and 16x lower latency over a sequential CNN+GRU+CLP edge GPU baseline, validated through iso-algorithm cross-platform benchmarking across three evaluation levels.

Autonomous driving perception demands accurate and efficient processing of three-dimensional sensor data under strict power constraints. Traditional convolutional neural networks achieve strong detection accuracy but are computationally intensive, limiting their suitability for deployment on resource-constrained neuromorphic platforms. Spiking neural networks offer a compelling alternative through event-driven sparse computation, yet their application to complex real-world perception tasks such as three-dimensional object detection remains limited. In this work, we propose an end-to-end spiking encoder-decoder network for object detection in bird's eye view representations of LiDAR point clouds, trained using surrogate gradient backpropagation. We train two variants: a membrane potential variant that reads continuous neuron state at the output stage for maximum accuracy, achieving $92.05$/$87.04$/$86.51$ AP at $\mathrm{IoU}\!=\!0.5$ (Easy/Moderate/Hard), and, a fully binary spiking variant that operates exclusively on spike trains at every layer for direct neuromorphic deployment. We evaluate four input spike encoding strategies and demonstrate that allowing the network to learn spike representations directly from data outperforms hand-crafted Poisson, latency, and z-axis encoding schemes on the KITTI benchmark, where sequential frames are unavailable and the BEV input is presented repeatedly across timesteps as a proxy for temporal streaming. A block-wise energy analysis demonstrates a $3.33\times$ reduction in synaptic operation energy over an equivalent CNN under conservative loop-based operation. Together, these results demonstrate the viability of spiking neural networks for accurate and energy-efficient neuromorphic perception in autonomous driving.

Deformable medical image registration aligns anatomical structures across images but remains computationally dense at 3D resolution. Spiking neural networks (SNNs) offer sparse event-driven computation, yet have not been systematically studied for deformable medical image registration. We introduce SpikeReg, a spiking U-Net for 3D brain MRI registration. SpikeReg is initialized from an analog ANN registration teacher, converted by layer-wise weight transfer and activation-percentile threshold calibration, and fine-tuned with a surrogate-gradient objective combining local cross-correlation, diffusion regularization, and spike-rate sparsity. On the OASIS Learn2Reg validation split ($19$ image pairs), SpikeReg reaches Dice $0.7474 \pm 0.032$, with no significant paired Dice difference from the ANN teacher ($0.7480 \pm 0.037$, $p = 0.67$), at a $12.8\%$ mean spike rate and a $55.5\times$ projected arithmetic-energy reduction under an event-sparse SynOps/MAC proxy relative to the dense-ANN baseline. We additionally report two negative findings: displacement distillation from the ANN teacher hurts performance, and ANN teachers trained with a label-Dice loss fail to transfer through rate-code conversion. Together these results show that dense geometric prediction can be performed under sparse event-driven computation, opening a path toward neuromorphic medical image registration.

Many-core neuromorphic systems accelerate Spiking Neural Networks (SNNs), yet their packet-based spike communication can spend substantial traffic and energy repeatedly transmitting destination addresses. This overhead is amplified by the small payload of spike packets: in representative workloads, duplicate address transmissions account for up to 49% of the total traffic. This paper presents UniSpike, a hardware-software co-design that removes address redundancy by aggregating spikes destined for the same core into compact packets. UniSpike combines destination-centric spike scheduling, lightweight runtime packet assembly hardware, and destination-aware SNN partitioning. Across diverse SNN workloads, UniSpike reduces traffic by 1.93$\times$ on average, delivering 1.77$\times$ speedup and 1.50$\times$ energy efficiency improvement over state-of-the-art designs.

Spiking Neural Networks (SNNs) provide an energy-efficient paradigm for visual recognition. We present SpikingMoE, which integrates a spike-driven Transformer with a Mixture-of-Experts (MoE) framework for dynamic computation. Inspired by the lateral geniculate nucleus (LGN), a spike-driven prompt (SDprompt) enables input-dependent expert routing in a biologically plausible manner. By replacing standard MLPs with spike-compatible expert modules and enforcing binary spike communication, SpikingMoE is designed for neuromorphic hardware. Experiments on CIFAR-10 and CIFAR-100 achieve 94.09% and 74.54% top-1 accuracy, showing that modular expert routing can be incorporated while retaining reasonable performance. To our knowledge, SpikingMoE is the first open-source SNN framework that integrates MoE into a spike-driven Transformer with LGN-inspired routing.

We investigate how internal representations emerge across hierarchical processing systems by introducing a neuroscience-inspired framework for analyzing deep spiking neural networks (SNN) through the lens of functional connectivity. Drawing on concepts from systems neuroscience and information theory, we form the first-order functionally-connected (1FC) group of a neuron based on its statistically significant pairwise correlations with neurons from the previous layer of a trained SNN architecture. We then track its response properties during inference under various conditions. Our analysis shows that several principles of functional connectivity previously observed in biological cortex are preserved in spiking ResNet architectures. These 1FC ensembles display interesting properties: their aggregate cofiring reliably predicts downstream neuronal responses through a robust, ReLU-like input-output relationship, whose gain scales systematically with ensemble size. Reliable encoding of the presented class emerges only during high 1FC cofiring events, which themselves occur infrequently, indicating that informative representations are concentrated in rare but highly coordinated activity patterns. Under uniform random noise or adversarial perturbations, these response profiles are disrupted, particularly in early and intermediate layers. This enables a targeted high-resolution interrogation at specific nodes and pathways. We showed that the functional connectivity structure is shaped by learning and this structure breaks under weight permutation. These establish 1FC ensembles as a functionally meaningful substrate for input encoding and information transfer, with potential implications in designing targeted fine-grained diagnostics on the information flow.

The Thousand Brains Theory (TBT) and its open-source Monty framework model object recognition through sensorimotor inference -- identifying objects by actively moving a sensor across their surface and building evidence contact by contact. The current implementation encodes each contact as a dense floating-point vector. While Monty tracks inter-step displacement and accumulates evidence across contacts, it treats the feature activation pattern at each contact as an unordered set - the directional sequence in which features are encountered carries no representational weight. In TBT, the sequence of contacts carries spatial meaning: knowing that feature A was felt before feature B during a left-to-right sweep tells you something about where A and B sit on the object. Dense vectors discard this ordering. We propose replacing dense vectors with rank-order spike packets: each contact produces a brief burst of neural events where the most strongly activated neuron fires first. The time gap between successive bursts implicitly encodes sensor displacement without explicit coordinate calculations. A biologically motivated learning rule (STDP) encodes traversal direction into synaptic weights. A learnable parameter lambda adjusts reliance on earlier versus recent contacts, adapting to each object's geometry. We derive three testable predictions and specify an implementation of four components in approximately 450 lines of NumPy. Three synthetic experiments confirm the core claims: temporal coding achieves perfect discrimination accuracy on objects with identical features in different spatial arrangements, where dense accumulation performs at chance; temporal coding maintains a 30-50 percentage point advantage across all tested noise levels; the adaptive lambda converges to distinct values, reflecting object geometric complexity. End-to-end evaluation on Monty's YCB benchmark is left for future work.

Natively trained spiking language models must preserve information across time while operating through sparse binary activations, a combination that has produced a persistent quality gap relative to dense Transformers. We present SymbolicLight V1, a spike-gated dual-path language model that couples binary Leaky Integrate-and-Fire (LIF) dynamics with a continuous residual stream. Its Dual-Path SparseTCAM mixer combines a first-order exponential-decay state with windowed local attention on the continuous residual stream, followed by a context-conditioned decoding head. We train four 194M-parameter models from scratch on a 3B-token, 10-domain Chinese-English corpus. On a fixed token-weighted evaluation set the runs reach perplexity (PPL) 8.88-8.93 (mean 8.904, sample standard deviation 0.019). Separation of this set from the training streams has not been verified. Training-time encoder probes have a mean zero-spike fraction of 89.96%; this is not a whole-model sparsity measure. Code tokens are 43.7% of that set; the unweighted mean of the ten domain PPLs is 29.38. Under the same corpus, tokenizer, token budget, and hardware, the token-weighted mean is 7.7% above GPT-2 201M (PPL 8.27). Across five zero-shot benchmarks the two 200M-scale models show no clear accuracy separation. Under sampling with temperature 0.7 and top-k 50, SymbolicLight produces lower 4-gram repetition; an entropy-modulated rule reverses that ranking. On an RTX 2080 Ti, measured generation throughput is 22.8 versus 91.5 token/s; post-generation power readings give rough energy estimates of 2,848 versus 905 mJ/token, without power integration over generation.

Spiking neural networks (SNNs) exploit event-driven and addition-only computation to substantially improve efficiency for intelligent computation. A key temporal property of SNNs, elastic inference, allows outputs to emerge progressively, enabling responses to salient inputs much earlier than full evaluation. However, existing SNN-specific accelerators cannot capitalize on this property. Layer-by-layer designs emit outputs only after all layers are complete, while time-step-by-time-step designs rely on coarse-grained, layer-wise pipelines that require synchronizing all spines/tokens within a layer. This barrier prevents results from being forwarded immediately, delaying the earliest possible response and forfeiting the benefits of elastic inference. To address these challenges, we propose ELSA, a near-SRAM dataflow architecture that realizes true elastic inference through a fine-grained spine/token-wise pipeline and hardware optimizations tailored to SNNs. ELSA forwards each spine/token immediately upon production, forming a continuous streaming pipeline that substantially reduces the latency to the first response. To enhance this lightweight execution, ELSA introduces a bundled address event representation protocol to lower communication traffic of network-on-chip (NoC), and leverages mini-batch spiking Gustavson-product to cut memory access and exploit inherent sparsity. Combined with mapping and scheduling optimizations, ELSA achieves efficient, event-driven computation without compromising accuracy. Experiments show that SNNs can outperform quantized artificial neural networks (QANNs) while maintaining on-par accuracy. For a 4-bit ResNet-50, ELSA achieves 3.4$\times$ speedup and 13.6$\times$ higher energy efficiency over the SOTA QANN accelerator (ANT), and 2.9$\times$ speedup and 22.1$\times$ energy efficiency gains over the SOTA SNN accelerator (PAICORE).

Adaptive robot navigation in dynamic environments requires policies that can reach the target reliably while producing efficient and stable trajectories. This paper presents Q-SpiRL, a quantum spiking reinforcement learning framework for obstacle-aware robot navigation. The framework develops and evaluates five agent families: tabular Q-learning, classical MLP, classical SNN, quantum-enhanced MLP (QMLP), and quantum-enhanced spiking neural network (QSNN). While all models are implemented under a unified training and evaluation pipeline, the QSNN is the central architecture of interest, as it combines spike-based temporal processing with variational quantum feature transformation. Experiments are conducted across three grid-world environments of increasing size, namely 20x20, 30x30, and 40x40, with both static and dynamic obstacles. Performance is assessed using success rate, success-weighted path length, path length, and turn rate under deterministic inference. Results show that QSNN achieves the strongest overall trade-off between task completion, trajectory efficiency, and motion smoothness, reaching up to 99% success rate while maintaining high path efficiency in the most challenging setting. Execution on IBM quantum hardware further demonstrates the feasibility of deploying the proposed hybrid policy under real-device conditions.

This work presents E-ReCON, a 16 Kb energy and resource-efficient digital compute-in-memory (DCIM) macro based on a compact 3T1R ReRAM bitcell for edge-AI inference. The proposed bitcell occupies only 0.85 um^2 and supports reliable AND-based in-memory multiplication for both conventional convolutional neural network (CNN) and spiking neural network (SNN) workloads. To reduce accumulation overhead, a novel interleaved 10T/28T adder tree is introduced, reducing transistor count and power consumption by 37% and 28%, respectively, compared to a conventional 28T RCA-based design. Implemented in 65 nm CMOS at 1.2 V, the proposed macro achieves a minimum latency of 0.48 ns, throughput of 2.31-3.1 TOPS, and energy efficiency of up to 419 TOPS/W. When evaluated on LeNet-5, AlexNet, and CNN-8 models, the macro achieves 97.81%, 93.23%, and 96.51% accuracy on MNIST/A-Z, CIFAR10, and SVHN datasets, respectively. In addition, 40% pruning preserves nearly 99.8% of the original accuracy while reducing MAC operations and computation cycles. For SNN-oriented workloads, the proposed AND-type bitcell efficiently supports spike-weight multiplication with low switching activity, where the 2A2W configuration achieves accuracy close to the FP32 baseline across VGG-8, VGG-16, and ResNet-18 networks on CIFAR-10, CIFAR-100, and ImageNet-1K datasets. Compared to prior ADC-based ReRAM-CIM designs, the proposed architecture improves latency and energy efficiency by nearly 30-40% while maintaining robust operation under full PVT and ReRAM variability. Overall, E-ReCON provides a scalable, low-latency, and energy-efficient nvCIM platform for next-generation edge-AI, IoT, biomedical sensing, and neuromorphic applications.

ANN-to-SNN conversion offers a practical, training-free route to spiking large language models. However, current pipelines primarily focus on spike-driven realizations for Transformer linear-algebra operations, while providing limited support for key nonlinear operators. This gap limits compatibility with neuromorphic-style execution constraints, where such nonlinearities typically require division, exponentiation, or norm computations that are not naturally supported by standard leaky integrate-and-fire dynamics. To solve this problem, we propose a plug-and-play framework that implements spike-friendly approximations for Transformer nonlinearities and integrates into existing ANN-to-SNN pipelines. Our method decomposes these nonlinear computations into three recurring primitives -- division, exponentiation, and $\ell_2$ norms -- and realizes them via population computation using LIF neuron groups, combined with lightweight bit-shift scaling to avoid floating-point arithmetic. By composing these primitives as modular operator blocks, our framework supports common Transformer nonlinearities (e.g., Softmax, SiLU, and normalization) without any fine-tuning. Experiments on a range of LLMs Transformers show that selectively replacing the targeted nonlinear operators incurs less than a $1\%$ accuracy drop across all evaluated tasks.

Flapping-Wing Micro Aerial Vehicles (FWMAVs) provide exceptional maneuverability and aerodynamic efficiency but pose significant challenges for onboard control due to nonlinear dynamics and stringent Size, Weight, and Power (SWaP) constraints, as exemplified by a butterfly-inspired robot less than 30 gram. To this end, we present a hierarchical neuromorphic control framework that enables fully onboard, closed-loop flight on a widely available, resource-constrained ESP32 microcontroller with a unit cost of approximately $5. Specifically, our method deploys two lightweight Spiking Neural Networks (SNNs) onboard: one for state estimation from raw sensory feedback and another for control via modulation of a Central Pattern Generator (CPG) for wing actuation. Trained by imitation learning, the system achieves stable pitch and heading angle tracking during untethered real-world flight. Experimental results further reveal that the SNN-based controller reduces latency by 36% (1059us to 680us) and power by 18% (0.033W to 0.027W) for inference compared to the conventional Artificial Neural Network (ANN) baseline, demonstrating the viability of spike-based computation without specialized hardware. To the best of our knowledge, this work constitutes the first demonstration of fully onboard neuromorphic control for autonomous flight of a FWMAV, highlighting the potential of SNNs to enable energy-efficient autonomy under stringent SWaP constraints. Visual abstract: http://bit.ly/4nI8ECY Code: https://anonymous.4open.science/r/Espikify-76E3/

Surrogate Safety Measures (SSMs) are extensively utilised in the evaluation of traffic risk in automated driving contexts. However, the majority of SSM-based evaluations employ fixed thresholds that fail to capture the human response to sustained borderline conditions or the reaction to brief, high-risk peaks. The present work proposes a biologically inspired reinterpretation of SSM thresholds. This is modelled as spiking thresholds of leaky integrate-and-fire (LIF) neurons, with multiple SSM inputs combined into a spiking neural network (SNN). The SNN is trained to emit spikes that are aligned with human braking onsets. The training data was recorded in a controlled car-following experiment using the 3D-CoAutoSim platform with CARLA/Unreal and a 6-DOF motion platform, where induced critical events were generated. The results demonstrate that the learned spiking activity qualitatively aligns with braking behaviour across scenarios and captures reactions that are not consistently explained by threshold crossings alone. Analysis across participants further indicates that learned input thresholds remain relatively consistent, while learned decay factors encode different temporal sensitivities for the SSMs. The findings of this study indicate that spiking dynamics may serve as a mechanism to facilitate the convergence of objective SSMs with subjective human safety perception.

Accurate and efficient time-series forecasting remains a challenging problem for both classical and quantum neural architectures, particularly in multivariate environmental settings. This work adapts the Quantum Leaky Integrate-and-Fire (QLIF) spiking neural network for time-series regression tasks, specifically short-term multivariate weather forecasting. We extend QLIF beyond classification and demonstrate its applicability to continuous-valued prediction problems. The QLIF-CAST model encodes neuron excitation states as single-qubit quantum superpositions, driven by R_x rotation gates and T1 relaxation decay, and is embedded within a hybrid quantum-classical recurrent architecture. We conduct two distinct evaluations. First, a controlled comparison against a parameter-matched classical LIF baseline on a multivariate weather dataset shows that QLIF-CAST achieves 15.4% lower MSE and 4.4% lower MAE, demonstrating that quantum neuronal dynamics reduce prediction error over classical equivalents. Second, a cross-domain comparative analysis with state-of-the-art quantum LSTM (QLSTM) and quantum neural network (QNN) models on air quality and wind speed benchmarks reveals that QLIF-CAST converges in up to 94% less training time, occupying a distinct position in the speed-error trade-off space. Hardware verification on IBM Marrakesh (156-qubit QPU) confirms reliable circuit execution with only 1.2% average deviation from simulation.

Deploying adaptive intelligence at the edge remains challenging due to the high computational and energy cost of training neural models. Spiking Neural Networks (SNNs) offer a promising alternative, but enabling on-device learning requires hardware-algorithm co-design. This paper presents SPIKER-LL, an FPGA-based SNN accelerator that extends the open-source Spiker+ inference architecture with efficient support for the STSF local learning rule. Through targeted microarchitectural extensions, SPIKER-LL performs inference and online learning with minimal overhead. Across MNIST, F-MNIST, and DIGITS, it achieves up to 93% accuracy, sub-millisecond latency, and less than 0.1 mJ per inference, while remaining DSP-free and highly scalable for edge-FPGA deployments.

Deploying event-camera object detectors is constrained by per-frame labeling requirements and GPU compute demands. This work introduces three local spike-timing-dependent plasticity (STDP) modules, including sequence, candidate, and tube-reliability modules, that operate on a single CPU thread without GPU support. On the FRED drone benchmark, the proposed framework spans three label-efficient supervision tiers. A strict zero-label detector achieves 53.8% mAP@30, approximately 26 train-derived bits achieve 76.9% mAP@30, and an STDP candidate-reliability gate achieves 78.60 +/- 0.42% mAP@30. Under acquisition-order drift, the cohort gate outperforms streaming k-means by 2.03 +/- 0.58 percentage points across 20 of 20 positive trials, while a no-drift control falsifies the effect. STDP reduces single-model variance by 6.6 times, and one trained gate matches a 44-seed ensemble bound. The gate transfers to Intel Lava with 89% top-2 agreement. On the EVUAV benchmark, a tube-level STDP layer reduces false alarms from 454 to 331e-4 at Pd >= 88%. Dense gradient-trained detectors cannot provide this combination of gradient training, dense matrix multiplication, and local plasticity-free operation by construction.

Von Economo neurons (VENs) are selectively lost in behavioural-variant frontotemporal dementia (bvFTD) and reduced in autism spectrum conditions (ASC), yet their computational role in social learning remains unexplained. We train a spiking neural network (the VENCircuit) embedding VEN-like projection neurons (K=40, 2% of total) in a recurrent pyramidal circuit across 50 matched random initialisations with and without VENs. The network is trained on a controlled binary classification task; we make no claim to model social cognition directly. VEN-intact networks converged in 49/50 cases (98%) versus 35/50 (70%) for VEN-ablated networks (Fisher's exact OR=21.0, 95% CI 2.7-167, p=8.7e-5). Failed ablated networks showed complete absence of learning, inconsistent with a speed-of-learning account. Phase-ablation experiments show VEN removal is most disruptive during mid-training (epochs 5-25), when a co-adaptive dependency forms in the pyramidal circuit. We derive a formal account showing VENs provide a direct gradient pathway immune to Jacobian instabilities affecting the recurrent circuit. Inference-time VEN ablation caused a significant performance drop (Wilcoxon p=0.022), ranging from no change (16/20 networks) to catastrophic collapse (0.989 to 0.620). VENs function as acquisition scaffolds whose developmental absence produces stochastic learning failure - a computational analogue of variable social skill acquisition in ASC - with falsifiable predictions for organoid and electrophysiology studies.

We propose a scalable neuromorphic architecture based on spiking dynamics emerging from the autonomous time-continuous evolution of clockless (asynchronous) digital circuits. Implemented on commercially available field-programmable gate arrays (FPGAs), our system implements networks of interacting Boolean spiking neurons with configurable excitatory and inhibitory synaptic weights. A complete processing pipeline enables efficient handling of spike-encoded data for solving machine-learning tasks. We demonstrate competitive performance for an audio classification task with spike-based encoding and high-speed processing. Power consumption is significantly lower than traditional digital implementations; this makes our approach an efficient alternative that bridges the gap to dedicated analog neuromorphic systems without the need for specialized hardware design. More generally, our approach establishes clockless digital hardware as a viable platform for neuromorphic computing. It paves the way for reconfigurable chips to be turned into energy-efficient quasi-analog neuromorphic processors.

Spiking neural networks (SNNs) hold promise for demonstrating superior learning and representation capabilities in deep models. Given the tremendous success of ResNet in deep learning, it would naturally follow to train deep SNNs with residual learning. However, existing residual structures for constructing deep SNNs still present challenges of spike redundancy or information loss, as well as redundant learning. In the present study, we first aim to address issues of relative spike redundancy in identity mapping and information loss in non-identity mapping. To this end, we propose an OR-ADD (OA) shortcut connection to merge output spikes/currents from two branches in the residual structure. Furthermore, to mitigate redundant learning in the backbone branch of the residual structure, we introduce the concept of XOR meta-residuals, i.e., selecting pre-learning residuals using the Exclusive-OR (XOR) operation for the backbone branch. Finally, by integrating the OA shortcut and XOR meta-residuals, we devise the XOR residual block and further construct XOResNet with varying depths based on this block. Extensive experiments on four datasets, Fashion-MNIST, CIFAR-10, CIFAR-100, and miniImageNet, show that the proposed XOResNet outperforms existing state-of-the-art deep SNNs optimized via gradient descent. These results validate the effectiveness of our OA shortcut and XOR meta-residual components in overcoming fundamental limitations of residual learning in SNNs, providing new architectural insights for building high-performance neuromorphic systems.

Spiking neural networks (SNNs) are biologically inspired energy-efficient models that use sparse binary spike-based communication between neurons, making them attractive for resource-constrained edge devices. Federated learning enables such devices to train collaboratively without sharing raw data. In time-series applications, edge devices often collect data at different time resolutions due to hardware and energy constraints. This temporal heterogeneity poses a fundamental challenge for federated learning: parameters learned at one temporal resolution do not necessarily transfer directly to another, which might result in the naive federated averaging being ineffective. Targeting SNNs and, more broadly, deep networks with stateful neurons, we propose a federated learning framework that addresses this temporal resolution mismatch. We investigate how neuron parameters learned from data at different temporal resolutions and model aggregation should be integrated. We evaluate the proposed framework across two SNN-native benchmark datasets (SHD and DVS-Gesture) under a range of resolution heterogeneity scenarios. Our results show that the proposed adaptation methods can substantially recover accuracy lost due to temporal mismatch, hence enabling each client to train at their local temporal resolution while remaining compatible with the global model.

The rapid expansion of spiking neural networks (SNNs) has led to a proliferation of training algorithms that differ widely in biological inspiration, computational structure, and hardware suitability. Despite this progress, the field lacks a unified, fine-grained taxonomy that systematically organizes these approaches and clarifies their conceptual relationships. This survey provides a comprehensive taxonomy of SNN training algorithms, spanning surrogate-gradient backpropagation, local and three-factor learning rules, biologically inspired plasticity mechanisms, ANN-to-SNN conversion pipelines, and non-standard optimization strategies. We analyze each class in terms of its computational principles, learning signals, and locality properties. To support reproducible research, we release NeuroTrain, an open-source snnTorch-based framework that implements a representative set of these algorithms within a unified, modular, and extendable framework, enabling consistent benchmarking across datasets, architectures, and training regimes. By consolidating fragmented literature and providing a reusable benchmarking framework, this survey identifies common patterns, highlights open challenges, and outlines promising directions for future work on scalable, efficient SNN training.

Spiking Neural Networks (SNNs) have emerged with promising energy-efficient property, yet a substantial performance gap persists compared to Artificial Neural Networks (ANNs). This gap stems from at least two key limitations: first, conventional spiking neurons offer limited information representation capacity, underutilizing the rich dynamics of membrane potentials; second, fixed surrogate gradient (SG) functions across time steps leads to imprecise gradient propagation, impeding effective direct training. To address these two challenges, we propose a new direct training algorithm with three core innovations: first, a circulate-firing spiking neuron model that enhances information representation capacity by leveraging membrane potentials more effectively; second, a time-step-wise learnable surrogate gradient function, enabling accurate gradient estimation during backpropagation; third, a positive-negative balanced loss function to achieve equilibrium between positive and negative membrane potentials and further boost SNN performance. Extensive experiments demonstrate that our methods achieve competitive performance across multiple datasets. Our methods can generalize seamlessly to advanced architectures of Transformer, consistently outperforming existing methods. Our work highlights the effectiveness of further harnessing intrinsic membrane dynamics of SNNs for performance improvement, and thus open a new avenue for advancing high-performance spiking neural architectures.

Spiking Neural Networks (SNNs) offer compelling energy efficiency on neuromorphic hardware, yet their training remains challenging because the discrete spike threshold is non-differentiable. Surrogate-gradient methods sidestep this by approximating the derivative, but they impose backpropagation infrastructure that is incompatible with on-chip learning. Evolution Strategies (\es) are a natural gradient-free alternative, yet their computational cost scales with the number of parameters, making them impractical for large weight matrices. We present a method for training SNNs using EGGROLL, a low-rank factorisation of ES perturbations that reduces per-generation memory from $\mathcal{O}(mn)$ to $\mathcal{O}(r(m{+}n))$. Combining EGGROLL with a Leaky Integrate-and-Fire SNN on N-MNIST, we demonstrate that gradient-free training achieves 79.21% test accuracy while reducing per-generation wall-clock time by 2.23$\times$ relative to full-rank ES. Our results demonstrate EGGROLL is viable for SNN training, with a clear accuracy-speed tradeoff, compatible with training on neuromorphic hardware without surrogate gradients.

Spiking neural networks (SNNs), which are brain-inspired and spike-driven, achieve high energy efficiency. However, a performance gap between SNNs and artificial neural networks (ANNs) still remains. Knowledge distillation (KD) is commonly adopted to improve SNN performance, but existing methods typically enforce uniform alignment across all timesteps, either from a teacher network or through inter-temporal self-distillation, implicitly assuming that per-timestep predictions should be treated equally. In practice, SNN predictions vary and evolve over time, and intermediate timesteps need not all be individually correct even when the final aggregated output is correct. Under such conditions, effective distillation should not force every timestep toward the same supervision target, but instead provide corrective guidance to erroneous timesteps while preserving useful temporal dynamics. To address this issue, we propose Selective Alignment Knowledge Distillation (SeAl-KD), which selectively aligns class-level and temporal knowledge by equalizing competing logits at erroneous timesteps and reweighting temporal alignment based on confidence and inter-timestep similarity. Extensive experiments on static image and neuromorphic event-based datasets demonstrate consistent improvements over existing distillation methods. The code is available at https://github.com/KaiSUN1/SeAl

Spiking Neural Networks (SNNs) are a promising framework for event-driven temporal processing. Prior work has improved temporal modeling through richer neuron dynamics and network-level mechanisms such as recurrence and delays, but it remains unclear how individual spiking neurons should specialize within a network. In this work, we introduce FiTS, a spiking neuron that factorizes temporal computation within each neuron into Frequency Selectivity (FS) and Temporal Shaping (TS). The FS module parameterizes each neuron's target frequency as the maximizer of its subthreshold magnitude response, while the TS module reshapes when frequency components contribute to membrane voltage accumulation through group-delay modulation. On auditory benchmarks where frequency selectivity and timing are central to the input structure, FiTS consistently improves over a plain Leaky Integrate-and-Fire (LIF) baseline in simple feedforward SNNs without recurrence or network-level delays, while remaining competitive with strong temporal SNN baselines. Beyond accuracy, the learned target frequencies and group-delay shifts provide interpretable neuron-level summaries of the frequency and timing organization learned within the network.